An interesting cloverleaf structure was proposed for the ori region with bound O protein in DNA Sequences and Structural Homologies of the Replication Origins of Lambdoid Bacteriophages, R. Grosschedl, H. Hobom, Nature 277, 621-627 (1979). What precisely does the DNAse footprinting of O bound to ori say about this model? See Purified Bacteriophage Lambda O Protein Binds to Four Repeating Sequences at the Lambda Replication Origin, T. Tsurimoto, K. Matsubara, Nucleic Acids Res. 9, 1789-1799 (1981).
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